1. Human Genetics and Cellular Genetics, Wellcome Trust Sanger Institute, Hinxton CB10 1SA, UK
2. Department of Oncology and Metabolism, University of Sheffield, Sheffield, S10 2RX, UK
3. Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford OX3 7BN, UK
4. Human Genetics, Wellcome Trust Sanger Institute, Hinxton CB10 1SA, UK