1. Department of Biological Sciences, University of Lethbridge , 4401 University Drive, Lethbridge, AB T1K 3M4 , Canada
2. Alberta Beekeepers Commission , 11434-168 Street #102, Edmonton, AB T5M 3T9 , Canada
3. National Bee Diagnostics Centre, Northwestern Polytechnic , PO Box 1118, Beaverlodge, AB T0H 0C0 , Canada