1. Key Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China
2. CAS Center for Excellence in Life and Paleoenvironment, Beijing 100044, China
3. Department of Biological Sciences, North Carolina State University, Raleigh, NC 27695, USA
4. Honors Program, Chapman University, Orange, CA 92866, USA
5. Department of Biology, Dalhousie University, Halifax, B3H 4R2, Canada
6. Department of Pathology and Anatomical Sciences, University of Missouri, Columbia, MO 65211, USA
7. North Carolina Museum of Natural Sciences, Raleigh, NC 27601, USA
8. Department of Geology, University of Lund, 22362, Sweden