1. School of Geographical Sciences, Northeast Normal University, Changchun, Jilin Province, China
2. Jilin Provincial Key Laboratory of Animal Resource Conservation and Utilization, Changchun, Jilin Province, China
3. Key Laboratory of Vegetation Ecology, Ministry of Education, Northeast Normal University, Changchun, Jilin Province, China
4. Department of Entomology and Plant Pathology, University of Tennessee, Knoxville, TN