1. College of Biological and Pharmaceutical Sciences, China Three Gorges University, Yichang, Hubei, China
2. Hubei International Scientific and Technological Cooperation Center of Ecological Conservation and Management in Three Gorges Area, China Three Gorges University, Yichang, China
3. Center for Global Change and Earth Observations/Geography, Environment and Spatial Sciences, Michigan State University, East Lansing, MI, USA
4. Key Laboratory of Aquatic Botany and Watershed Ecology, Wuhan Botanical Garden, the Chinese Academy of Sciences, Wuhan, China