1. Laboratory of Isotope Biogeochemistry, Bayreuth Center of Ecology and Environmental (BayCEER), University of Bayreuth , Universitätsstrasse 30, 95440 Bayreuth , Germany
2. Yunnan Laboratory for Conservation of Rare, Endangered & Endemic Forest Plants, National Forestry and Grassland Administration, Yunnan Academy of Forestry and Grassland , Kunming 650201, Yunnan , People’s Republic of China
3. Department of Life Science, National Taiwan University , Taipei 10617 , Taiwan
4. Institute of Ecology and Evolutionary Biology, National Taiwan University , Taipei 10617 , Taiwan