1. Department of Botany, Gombe State University, PMB 127, Tudun Wada , Gombe, Gombe State , Nigeria
2. Bolus Herbarium, Department of Biological Sciences, University of Cape Town , Private Bag X3, Rondebosch, Cape Town 7701 , South Africa
3. Department of Organismic and Evolutionary Biology, Harvard University Herbaria , 22 Divinity Avenue, Cambridge, MA 02138 , USA
4. Department of Botany and Zoology, Faculty of Science, Masaryk University , Kotlarska 2, 611 37 Brno , Czech Republic
5. Organismic Botany and Mycology, Institute of Plant Science and Microbiology, University of Hamburg , Germany