1. Armauer Hansen Research Institute, Addis Ababa, Ethiopia; Addis Ababa University, Addis Ababa, Hawassa University, Hawassa, Ethiopia
2. Addis Ababa University, Addis Ababa, Ethiopia
3. Armauer Hansen Research Institute, Addis Ababa, Ethiopia
4. REACH Ethiopia, Hawassa, Ethiopia, Liverpool School of Tropical Medicine, Liverpool, UK
5. Emory University School of Medicine, Atlanta, Georgia, USA