1. Graduate School of Agriculture, Kyoto University; Kitashirakawa Oiwakecho Sakyo-ku, Kyoto 606-8502 Japan
2. Graduate School of Global Environmental Studies, Kyoto Univ.; Yoshida Honmachi Sakyo-ku, Kyoto 606-8501 Japan
3. Japan International Research Center for Agricultural Sciences (JIRCAS); 1-1 Ohwashi Tsukuba Ibaraki 305-8686 Japan
4. Graduate School of Urban Environmental Sciences, Tokyo Metropolitan University; 2-2 Minami-Osawa Hachioji Tokyo 192-0364 Japan