1. School of Nature Conservation, Beijing Forestry University 100083, P. R. China
2. Ecologie Systématique Evolution, Université Paris-Sud, CNRS, AgroParisTech, Université Paris-Saclay, F-91405 Orsay Cedex, France
3. College of Life Science, Guizhou University, Guiyang 550025, P. R. China
4. Shaanxi Hanzhong Crested Ibis National Nature Reserve, Shaanxi 723300, P. R. China
5. Department of Biological Sciences, Purdue University, West Lafayette, Indiana, USA