1. Institute of Grassland and Environmental Research, Plas Gogerddan, Aberystwyth, Ceredigion, SY23 3EB, UK
2. Biodiversity Department, Kenya Wildlife Service, PO Box 40241, Nairobi, Kenya
3. Department of Agriculture, The University of Reading, Earley Gate, PO Box 236, Reading, Berkshire, RG6 6AT, UK