1. School of Medicine, Cardiff University 1 Division of Infection and Immunity, , Cardiff, UK
2. Systems Immunity Research Institute, Cardiff University 2 , Cardiff, UK
3. Lydia Becker Institute of Immunology and Inflammation, Immunity to Infection and Respiratory Medicine, School of Biological Sciences, Faculty of Biology, Medicine and Health, University of Manchester 3 Division of Immunology, , Manchester, UK