1. Department of Molecular Pharmacology, Tokyo Medical and Dental University
2. University of Tokyo, Tokyo 113-8655, Japan
3. Ehime University, Ehime 791-0295, Japan
4. Center of Excellence Program for Frontier Research on Molecular Destruction and Reconstruction of Tooth and Bone, Core to Core Program for Advanced Bone and Joint Science, Japan Society for Promotion of Science, Tokyo 101-0062, Japan
5. Integrated Action Initiative, Core to Core Program for Advanced Bone and Joint Science, Japan Society for Promotion of Science, Tokyo 101-0062, Japan