1. Ludwig Institute for Cancer Research, PO Royal Melbourne Hospital, VIC 3050, Australia
2. Department of Pathology, Peter MacCallum Institute, East Melbourne VIC 3002, Australia
3. The Walter and Eliza Hall Institute, PO Royal Melbourne Hospital, VIC 3050, Australia
4. Department of Pathology, University of Sydney, Sydney, NSW 2006, Australia
5. Department of Surgery, The University of Melbourne Parkville, VIC 3052, Australia