1. Graduate School of Science and Technology, Shizuoka University, Hamamatsu 432-8561, Japan
2. Department of Marine Biosciences, Faculty of Marine Science, Tokyo University of Marine Science and Technology, Minato, Tokyo 108-8477, Japan
3. Department of Systems Engineering, Faculty of Engineering, Shizuoka University, Hamamatsu 432-8561, Japan
4. Department of Environmental and Forest Biology, State University of New York College of Environmental Science and Forestry, Syracuse, NY 13210, U.S.A.