1. Wissenschaftskolleg zu Berlin, Wallotstraße 19, Berlin, Germany
2. Evolutionary Biology, Universität Bielefeld, Konsequez 45, 33615 Bielefeld, Germany
3. Department of Biology, University of Florida, Gainesville, Florida 32611
4. Department of Biological Sciences, Macquarie University, North Ryde, New South Wales 2109, Australia
5. Organismal and Evolutionary Biology Research Program, Faculty of Biological and Environmental Sciences, Helsinki University, Helsinki, Finland; and Department of Biological and Environmental Science, University of Jyväskylä, Jyväskylä FI-40014, Finland