1. Wyoming Natural Diversity Database and Department of Zoology and Physiology, University of Wyoming, 1000 East University Avenue, Laramie, Wyoming 82071 USA
2. Department of Ecology, Montana State University, 310 Lewis Hall, Bozeman, Montana 59717 USA
3. Montana Cooperative Fishery Research Unit, Montana State University, P.O. Box 173460, Bozeman, Montana 59717 USA
4. United States Geological Survey, Montana Cooperative Fishery Research Unit, Montana State University, P.O. Box 173460, Bozeman, Montana 59717 USA
5. Native Fish Conservation Program, Yellowstone National Park, P.O. Box 168, Yellowstone National Park, Wyoming 82190 USA