1. Ecology and Environmental Sciences Program, School of Biology and Ecology, University of Maine, Orono, Maine 04469
2. Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, Michigan 48109
3. Department of Ecology and Evolutionary Biology, University of California, Santa Cruz, California 95064
4. Department of Ecology and Evolutionary Biology, University of California, Irvine, California 92697