1. Meliponini and Apini Research Laboratory, Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand
2. Department of Plant and Soil Science, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand
3. Science and Technology Research Institute, Chiang Mai University, Chiang Mai 50200, Thailand
4. Department of Plant Medicals, Andong National University, Andong 36729, Republic of Korea
5. Department of Horticulture, Oregon State University, Corvallis, OR 97331, USA