1. Research Center for Sub-tropical Fisheries, Seikai National Fisheries Research Institute, Japan Fisheries Research and Education Agency, Ishigaki, Okinawa, Japan
2. Research Center for Marine Invertebrates, National Research Institute of Fisheries and Environment of Inland Sea, Japan Fisheries Research and Education Agency, Onomichi, Hiroshima, Japan
3. Graduate School of Fisheries and Environmental Sciences, Nagasaki University, Nagasaki, Nagasaki, Japan