1. Institute Biology Leiden, Leiden University, Leiden, The Netherlands
2. Naturalis Biodiversity Center, Leiden, The Netherlands
3. Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, Kota Kinabalu, Sabah, Malaysia
4. Centrum Wiskunde & Informatica, Science Park, Amsterdam, The Netherlands
5. University of California San Francisco (UCSF), Anatomy Department, Genentech Hall, San Francisco, CA, United States