1. Department of Ecology and Evolution, University of Lausanne, Biophore, Lausanne, Switzerland
2. Instituto Pirenaico de Ecología (IPE-CSIC), Jaca, Spain
3. Fundación Araid, Zaragoza, Spain
4. Department of Biodiversity and Evolutionary Biology, Museo Nacional de Ciencias Naturales (MNCN-CSIC), Madrid, Spain