1. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China
2. CAS Center for Excellence in Life and Paleoenvironment, Beijing, China
3. University of Chinese Academy of Sciences, Beijing, China
4. Nanjing Normal University, Nanjing, Jiangsu, China
5. Department of Vertebrate Paleontology, Carnegie Museum of Natural History, Pittsburgh, PA, United States of America
6. School of Dentistry, University of California, Los Angeles, Los Angeles, CA, United States of America
7. Natural History Museum of Los Angeles County, Los Angeles, CA, United States of America