1. Department of Natural Resource Ecology and Management, Iowa State University, Ames, IA, United States of America
2. Department of Natural and Environmental Sciences, Western State Colorado University, Gunnison, CO, United States of America
3. Department of Veterinary Microbiology and Preventive Medicine, Iowa State University, Ames, IA, United States of America
4. Animal and Plant Health Inspection Service, Wildlife Services, United States Department of Agriculture, Urbandale, IA, United States of America
5. Department of Veterinary Diagnostic and Production Animal Medicine, Iowa State University, Ames, IA, United States of America