1. Biodiversity Division, National Institute for Environmental Studies, Tsukuba, Ibaraki, Japan
2. University of Tsukuba, Tsukuba, Ibaraki, Japan
3. Fukushima Regional Collaborative Research Center, National Institute for Environmental Studies, Miharu, Fukushima, Japan
4. Unaffiliated, Tsukuba, Ibaraki, Japan
5. National Parks Awareness Center, Edogawa University, Nagareyama, Chiba, Japan