1. State Key Laboratory of Grassland Agro-ecosystems; Key Laboratory of Grassland Livestock Industry Innovation, Ministry of Agriculture and Rural Affairs; Engineering Research Center of Grassland Industry, Ministry of Education; Gansu Tech Innovation Centre, Lanzhou University, Lanzhou, Gansu, China
2. Key Laboratory of Medicinal Plant and Animal Resources of the Qinghai-Tibetan Plateau, School of Life Science, Qinghai Normal University, Xining, Qinghai, China
3. Department of Plant Biology, Rutgers University, New Brunswick, NJ, United States of America
4. Grasslands Research Centre, AgResearch Limited, Palmerston North, New Zealand