1. Cibio, Research Centre in Biodiversity and Genetic Resources, InBio, Universidade do Porto, Campus Agrário de Vairão, Rua Padre Armando Quintas, Portugal, Porto, Portugal
2. Department of Biology, University of Massachussetts Boston, Boston, MA, USA
3. Zoological Institute, Braunschweig University of Technology, Mendelssohnstr. 4, Germany, Braunschweig, Germany
4. Madagascar Fauna and Flora Group, BP 442, 501 Toamasina, Madagascar, Toamasina, Madagascar
5. Max Planck Institute for Evolutionary Biology, August-Thienemann-Str. 2, Germany, Plön, Germany
6. Amphibian Survival Alliance c/o Durrell Wildlife Conservation Trust, Madagascar Programme, Lot II Y 49 J 12 Ampasanimalo, BP 8511 101 Antananarivo, Madagascar, Antananarivo, Madagascar