1. Key Laboratory of Tropical Marine Bio-resources and Ecology, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, Guangdong, PR China
2. College of Ocean and Earth Sciences, Xiamen University, Xiamen, Fujian, PR China
3. Hainan Key Laboratory for Sustainable Utilization of Tropical Bioresource, Hainan University, Haikou, Hainan, PR China