1. Tribhuvan University, Institute of Forestry, Kathmandu, Nepal
2. Tribhuvan University, Institute of Forestry, Pokhara, Nepal
3. Department of Forest Sciences, Faculty of Agriculture and Forestry, University of Helsinki, Helsinki, Finland
4. Department of Biological Sciences, University of Alberta, Edmonton, Canada
5. Department of Biology, University of Kentucky, Lexington, KY, United States of America
6. Department of Biology, Western Kentucky University, Bowling Green, KY, United States of America