1. School of Animal and Veterinary Sciences, The University of Adelaide, Roseworthy, South Australia, Australia
2. Department of Wildlife Studies, Faculty of Veterinary of Science, University of Pretoria, Onderstepoort, Gauteng, South Africa
3. Department of Veterinary Tropical Diseases, Hans Hoheisen Wildlife Research Station, University of Pretoria, Onderstepoort, Gauteng, South Africa
4. Mpumalanga Tourism and Parks Agency, Nelspruit, Mpumalanga, South Africa
5. Department of Biological Sciences, University of Venda, Thohoyandou, Limpopo, South Africa