1. South China Botanical Garden, Chinese Academy of Sciences, Guangzhou, Guangdong, China
2. Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Guangdong Institute of Applied Biological Resources, Guangzhou, Guangdong, China
3. University of Chinese Academy of Sciences, Beijing, China
4. Guangdong Luokeng Shinisaurus crocodilurus National Nature Reserve, Shaoguan, Guangdong, China
5. Guangxi Key Laboratory of Rare and Endangered Animal Ecology, Guangxi Normal University, Guilin, Guangxi, China