1. School of Natural Sciences, Massey University, Palmerston North, Manawatu, New Zealand
2. Molecular Epidemiology and Veterinary Public Health Laboratory—Hopkirk Research Institute, School of Veterinary Science, Massey University, Palmerston North, Manawatu, New Zealand
3. Centre for Tropical Water and Aquatic Ecosystem Research (TropWATER), James Cook University of North Queensland, Townsville, Queensland, Australia