1. Illinois Natural History Survey, Prairie Research Institute, University of Illinois at Urbana-Champaign, Champaign, IL, United States of America
2. Department of Natural Resources and Environmental Sciences, College of Agricultural, Consumer and Environmental Sciences, University of Illinois at Urbana-Champaign, Urbana, IL, United States of America
3. Program in Ecology, Evolution & Conservation Biology, University of Illinois at Urbana-Champaign, Urbana, IL, United States of America