1. Applied Biodiversity Research Division, South African National Biodiversity Institute, Cape Town, South Africa
2. Department of Zoology, University of Cape Town, Rondebosch, South Africa
3. Department of Botany and Zoology, University of Stellenbosch, Matieland, South Africa
4. DST-NRF Centre for Invasion Biology (CIB), Department of Zoology, Nelson Mandela Metropolitan University, Port Elizabeth, South Africa