1. Department of Geography and the Environment, University of Aberdeen, St Mary's, Elphinstone Road, Aberdeen AB24 3 UF, UK; School of Geography, University of Leeds, Leeds LS2 9JT, UK;
2. Department of Zoology, University of Cambridge, Downing Street, Cambridge CB2 3EJ, UK
3. School of Geography, University of Leeds, Leeds LS2 9JTj UK
4. NERC Radiocarbon Laboratory, Scottish Enterprise Technology Park, Rankine Avenue, East Kilbride G75 OQF, UK
5. Department of Geology, University of Patras, GR-26110 Rio-Patras, Greece