1. Department of Quaternary Geology, Lund University, Sölvegatan 12, SE-223 62 Lund, Sweden;
2. Department of Ecology, Evolution and Behaviour, University of M innesota, 1987 Upper Buford Circle, St Paul, M N 55108-6097, USA
3. Department of Biology and Environmental Sciences, Kalmar University, SE-391 82 Kalmar, Sweden