1. Department of Forest Resources, Ehime University, 3-5-7 Tarumi, Matsuyama 790-8566, Japan
2. Department of Biology and Chemistry, Växjö University, SE-351 95 Växjö, Sweden; Department of Quaternary Geology, Lund University, SE-223 63 Lund, Sweden
3. Department of Quaternary Geology, Lund University, SE-223 63 Lund, Sweden