1. State Key Laboratory of Desert and Oasis Ecology, Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences, Urümqi 830011, China
2. College of Life Sciences, Xinjiang Agricultural University, Urümqi 830052, China
3. Shangdong Provincial Key Laboratory of Water and Soil Conservation and Environmental Protection, College of Resources and Environment, Linyi University, Linyi 276000, China