1. Department of Agroecology, Aarhus University, DK-4200, Slagelse, Denmark
2. Department of Science and Conservation, National Tropical Botanical Garden, Kalāheo, HI, 96741, USA; Natural History Museum of Denmark, University of Copenhagen, DK-2100, Copenhagen, Denmark
3. School of Agriculture and Environment (PN433), Massey University, Palmerston North 4442, New Zealand