1. School of Life Science, Yunnan Normal University, Kunming, 650500, Yunnan, China; Key Laboratory of Yunnan for Biomass Energy and Biotechnology of Environment, Yunnan Normal University, Kunming, 650500 China
2. School of Life Science, Yunnan Normal University, Kunming, 650500, Yunnan, China
3. Faculty of Geography, Yunnan Normal University, Kunming 650500, Yunnan, China
4. Department of Biology, University of Kentucky, Lexington, Kentucky 40506-0225, USA
5. Department of Biology, University of Kentucky, Lexington, Kentucky 40506-0225, USA; Department of Plant and Soil Sciences, University of Kentucky, Lexington, Kentucky 40546-0312, USA