1. Tsukuba Primate Research Center, National Institutes of Biomedical Innovation, Health and Nutrition, Hachimandai 1-1, Tsukuba, Ibaraki 305-0843, Japan
2. College of Bioresource Sciences, Nihon University, Japan
3. The Corporation for Production and Research of Laboratory Primates, Japan
4. Mie University Graduate School of Medicine, Department of Molecular and Experimental Medicine, Japan