1. Center for Experimental Medicine and Systems Biology, The Institute of Medical Science, The University of Tokyo, 4-6-1, Shirokanedai, Minato-ku, Tokyo 108-8639, Japan
2. Department of Biophysics and Biochemistry, Graduate School of Science, The University of Tokyo, Tokyo 113-0032, Japan
3. Research Institute for Biomedical Sciences, Tokyo University of Science, Chiba 278-0022, Japan
4. Present address: Department of Biomedical Science, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo 113-8657, Japan
5. Present address: Department of Molecular Immunology, Medical Mycology Research Center, Chiba University, Chiba 263-8522, Japan