1. NIBS Laboratory Animal Research Station, Nippon Institute for Biological Science, 3331-114 Kamisasao, Kobuchisawa, Hokuto, Yamanashi 408-0041, Japan
2. Headquarters, Nippon Institute for Biological Science, 9-2221-1 Shinmachi, Ome, Tokyo 198-0024, Japan
3. Department of Molecular Pathology, Interdisciplinary Graduate School of Medicine and Engineering, University of Yamanashi, 1110 Shimokato, Chuo, Yamanashi 409-3898, Japan
4. Department of Biochemistry and Molecular Biology, Graduate School of Medical and Dental Sciences, Kagoshima University, 8-35-1 Sakuragaoka, Kagoshima-city, Kagoshima 890-8520, Japan