1. Division of Bio-resources, Department of Biotechnology, Frontier Science Research Center, University of Miyazaki, 5200 Kihara, Kiyotake, Miyazaki 889–1692, Japan
2. Present Address: WDB EUREKA Co., Ltd., 2-3-2 Marunouchi, Chiyoda-ku, Tokyo, Japan
3. Present Address: Faculty of Law and Economics, Okinawa University, 555 Kokuba, Naha, Okinawa 902-8521, Japan
4. Faculty of Science, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903-0213, Japan