1. Department of Cell and Developmental Biology, Vanderbilt University, Nashville, TN 37240, USA
2. Department of Molecular and Cellular Biochemistry, University of Kentucky, Lexington, KY 40536, USA
3. Markey Cancer Center, University of Kentucky, Lexington, KY 40536, USA
4. Key Laboratory of Bio-Resource and Eco-Environment of Ministry of Education, Sichuan University, Chengdu 610000, Sichuan, P. R. China
5. Department of Neuroscience, University of Kentucky, Lexington, KY 40536, USA