1. University of Michigan, College of Literature, Science, and The Arts, Ann Arbor, MI 48109, USA
2. Calico Life Sciences, South San Francisco, CA 94080, USA
3. Current Affiliation: WuXi AppTec, Shanghai, China
4. University of Michigan, Department of Pathology, Ann Arbor, MI 48109, USA
5. University of Michigan Geriatrics Center, Ann Arbor, MI 48109, USA