1. College of Animal Science, South China Agricultural University, Guangzhou, 510642, P.R. China
2. Guangdong Provincial Key Lab of Agro-Animal Genomics and Molecular Breeding, Guangzhou, 510642, P.R. China
3. Key Laboratory of Animal Health Aquaculture and Environmental Control, Guangdong, Guangzhou, 510642, P.R. China
4. South China Collaborative Innovation Center for Poultry Disease Control and Product Safety, Guangzhou, 510642, P.R. China
5. College of Science and Engineering, Jinan University, Guangzhou, 510632, P.R. China
6. USDA, Agriculture Research Service, Avian Disease and Oncology Laboratory, East Lansing, MI, 48823, USA