1. Department of Organismal Systems and Bioresilience, Biological Systems and Engineering Division, Lawrence Berkeley National Laboratory, Berkeley, CA 94720, USA
2. Present address: Department of Dermatology, University of Texas, Southwestern Medical Center, Dallas, TX 75390, USA
3. Department of Molecular Medicine, Institute of Basic Medical Science, University of Oslo, 0317 Oslo, Norway
4. Department of Pathology and Laboratory Medicine, University of California, Davis, CA 95817, USA
5. Department of Environmental and Radiological Health Sciences, Colorado State University, Fort Collins, CO 80523, USA