1. Department of Biology, University of Washington, Seattle, WA 98195, USA
2. Department of Pathology, University of Washington, Seattle, WA 98195, USA
3. School of Environmental and Forest Sciences, University of Washington, Seattle, WA 98195, USA
4. Wildlife Conservation Society, Bronx, NY 10460, USA
5. Global Penguin Society, University of Washington, Seattle, WA 98195, USA