1. Department of Ophthalmology, Keck School of Medicine of the University of Southern California, Los Angeles, CA 90089, USA
2. Arnold and Mabel Beckman Macular Research Center at the Doheny Eye Institute, Los Angeles, CA 90033, USA
3. Department of Ophthalmology, Kagoshima University, Graduate School of Medical and Dental Sciences, Kagoshima, Japan
4. These two authors contributed equally to this work
5. Department of Pathology, Keck School of Medicine of the University of Southern California, Los Angeles, CA 90089, USA